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A Sunny Day in Glasgow began as a collaboration between friends Ben Daniels and Ever Nalens, both of whom had recently returned to Philadelphia after several years in the UK. The band name came from Nalens, who had been living in Glasgow, and Daniels kept it after Nalens left the project. In 2006, Daniels asked his twin sisters, Robin and Lauren, to start singing on the songs they had been working on. In March of that year the trio self-released The Sunniest Day Ever EP to great acclaim from college radio stations and blogs. Their debut release, Scribble Mural Comic Journal, came out February 2007 and received favorable reviews from Pitchfork and Drowned in Sound among others.
2009 saw the band release their second studio LP, Ashes Grammar. During the recording process, the band was forced to undergo several member changes. The first of these changes came when former bassist Brice Hickey fell when loading equipment into his car, breaking several bones in his legs. This left Ben Daniels to record the bass parts for the album, but also meant that vocalist Robin Daniels had to leave to care for the bedridden Hickey, who is her boyfriend. Lauren Daniels also had to leave the group, due to attending grad school in Colorado, forcing Ben Daniels and guitarist/drummer Josh Meakim to recruit vocalist Annie Fredrickson, also a classically trained cellist and pianist. After the album was recorded, Ryan Newmyer joined as bassist and Meakim's friend Adam Herndon as drummer. In addition, through an advert via their Myspace blog, the band found another vocalist, Jen Goma. The new line-up completed all of the songs from the Ashes sessions and released them as the Nitetime Rainbows EP in March 2010 and "Autumn, Again" later in October.
On June 24, 2014 they released their album Sea When Absent.
